Class: Charophyceae

Perhaps the most differentiating aspect from the other 2 classes is that the nuclear membrane disappears before metaphase. and the spindle persists until after partitioning is complete

In some species furrowing occurs and in others a phragmoplast and cell plate is formed

Species produce a dormant zygote that undergoes zygotic meiosis so that all multicellular stages are haploid; in higher speices there is a clear case of oogamy = gametes differentiate into large eggs and smaller sperm.

Species are generally freshwater

Flagellated species are inserted laterally and and attach to a multilayered structure of microtubules so that the cells are asymmetric.


The order which we'll further explore in lab is Charales which includes Chara and Nitella.

 

Chara is quite intersting as it is quite advanced with an egg protected by secondary sexual cells.

 

from: http://vis-pc.plantbio.ohiou.edu/algaeimage/pages/Chara.html


Zygnemataceae - there is argument here where this group belongs - your text places in the Ulvophyceae but other sources list it here....they are known as the conjugating algae as two filaments will meet and form conjugation tubes.See lab notes for more information...

Spirogyra is another well known genera in this group.

 

 

Spirogyra


 

Desmids;can be quite beautiful; symmetrical 1-4+ celled floating phytoplankton.

 


 

 

Coleochaetales

The fifteen species of Coleochaetales have gained attention recently because they are implicated as the closest living relatives of the Plantae. Some species have parenchymatous growth, nurturing of the zygotes, and a fossil record of an appropriate age, all of which strengthen the implication

 

 

Coleochaete is a small genus of filamentous or parenchymatous green algae. The ca. five species occur commonly in freshwater environments worldwide, where they usually are found as epiphytes on aquatic plants. Coleochaete, and the allied genus Chaetosphaeridium, are thought to be the closest living algal relatives of the stoneworts

Individual vegetative cells of Coleochaete species are usually 5-20 µm in diameter and 1-4 times as long as broad, with a conspicuous cell wall. The cells contain a single plastid that occupies most of the cell periphery and contains at least one pyrenoid. (Chara), brittleworts (Nitella), and land plants.

The cells are organized into branched filaments that, together, produce a plant body less than 2 mm in diameter. This plant body is prostrate on, or embedded in, the substrate, with no plastid-bearing "erect" filaments extending from the substrate plane. In some species, the filaments are laterally compressed, forming a circular thallus with no apparent gaps between cells. Cultured plants that do not become fixed to a substrate may form spherical balls of cells.